Spanish journal of palaeontology. 2019. Vol. 34, no. 2

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    Primeros conodontos del Frasniense en el Complejo Maláguide de Granada (Cordillera Bética)
    (2021) Cabrera-Porras, Angela; Rodríguez-Cañero, Rosario; Martín-Algarra, Agustín
    Un delgado nivel calc.reo, localizado en la unidad tectónica más alta del Complejo Maláguide en la provincia de Granada, ha proporcionado una abundante fauna de conodontos (808 elementos por kg de muestra) integrada por Ancyrodella binodosa, Ancyrodella pristina, Icriodus difficilis, Icriodus symmetricus, Polygnathus decorosus, Polygnathus dengleri y Polygnathus dubius, entre otras especies. Esta asociación corresponde a una biofacies de polygnátidos y ha permitido asignar este nivel a las zonas MN1 y MN2 (Frasniense inicial), cuya presencia ha sido constatada por vez primera en el Complejo Maláguide. Esta es, asimismo, la primera datación del Frasniense en el sector central de la Cordillera Bética. El nivel se intercala entre pelitas con niveles areniscosos, volcanoclásticos y otros delgados horizontes carbonatados del Emsiense y del Fameniense, que yacen bajo la Fm. Falcoña (Carbonífero Inferior). Esta sucesión pre-Falcoña se encuentra en una unidad tectónica diferente a la que incluye la Fm. Santi Petri, con la que se correlaciona, aunque se depositó en ambientes menos distales y menos profundos que dicha formación, que es la más extensamente representada y característica del Devónico maláguide.
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    Palaeoentomological study of the lacustrine oil-shales of the lower Miocene San Chils locality (Ribesalbes-Alcora Basin, Castellón province, Spain)
    (2021) Álvarez-Parra, Sergio; Peñalver Mollá, Enrique
    For the first time, the study of the fossil record present in laminated bituminous dolostones from the San Chils locality, lower Miocene in age (ca. 19 Ma), located at the Ribesalbes-Alcora Basin, is addressed. The identification of 11 families and three genera belonging to six insect orders (Orthoptera, Thysanoptera, Hemiptera, Coleoptera, Hymenoptera and Diptera) has been carried out. Furthermore, ephippia of ladoceran crustaceans, one bird feather, vertebrate coprolites and plant remains have been found. The San Chils palaeodiversity has been compared to that of La Rinconada, a similar outcrop in the same basin and depositional unit. Both localities correspond to lacustrine deposits and were mined during the past two centuries. This work has increased the palaeontological richness of the Ribesalbes-Alcora Basin with new fi ndings such as specimens of the subfamily Corixinae (Hemiptera: Corixidae), some ant workers (Hymenoptera: ormicidae), a specimen assigned to the genus Exechia (Diptera: Mycetophilidae), ephippia of Daphnia  (Ctenodapnia ) sp. (Branchiopoda: Cladocera), the possible first report of the genus Sciocoris  (Hemiptera: Pentatomidae) for the Neogene fossil record and the possible presence of the family Aeolothripidae (Thysanoptera). The current collection of fossil specimens from San Chils is small, but shows that it shared similar palaeoenvironmental conditions to that of La Rinconada, both in the lake waters and in the surroundings. 
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    Estudio paleontológico de los gasterópodos y escafópodos (Mollusca) del Tortoniense superior de Arroyo Trujillo, Cantillana (Sevilla)
    (2021) Cárdenas, Joaquín; Bajo, Ildefonso; Maestre, M. Vicente
    Se ha estudiado el conjunto de gaster.podos y escafápodos correspondiente al Tortoniense superior en el yacimiento de Arroyo Trujillo (Cantillana, Sevilla), en el borde norte de la Cuenca del Guadalquivir. Se han identificado 194 taxones de gasterópodos y 8 de escafópodos. Todos ellos han sido localizados en los tramos III y IV, de los cinco diferenciados en el yacimiento (Cárdenas et al., 2017). De sus improntas tafonómicas se deduce que los ejemplares han sufrido arrastres puntuales, remociones junto con los sedimentos del fondo marino con posteriores resedimentaciones, y por sus escasas bioerosiones y bioincrustaciones, producidas por otros seres vivos, un enterramiento temprano. Se analiza la distribución espacial de las especies durante el Neógeno en los dominios Atlántico, Mediterráneo y Paratethys, y se constata una mayor coincidencia del conjunto de gasterópodos y escafópodos del yacimiento con el conjunto de éstos citados en la zona mediterránea que con los citados en la zona atlántica. También se analizan determinados movimientos migratorios de algunas especies entre los diferentes dominios marinos. Caso muy problemático e incierto es la dispersión de Turritella incisaeformis.  En cuanto a las condiciones ambientales, se deduce una batimetría de 20-40 m. para el tramo III y 50-80 m. para el tramo IV; una salinidad normal y temperatura tropical o subtropical corroborada por la abundancia de especies de gasterópodos termófilos.
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    Middle Eocene calcareous nannofossils in the Jaca transect (South-central Pyrenees Eocene Basin, Aragón river valley, Huesca)
    (2021) González-Lanchas, Alba; Remacha, Eduard; Oms, Oriol; Sierro, Francisco J.; Flores, Jose Abel
    The calcareous nanoplankton is studied from the base of the turbidite systems of the Upper Hecho Group (Jaca Basin, middle Eocene of the South-central Pyrenean Basin) up to the Gracionepel instabilities, within the Larrés slope Marls. This new chronostratigraphic contribution is of crucial importance for the improvement of the detailed temporal and spatial correlation framework of the genetically related depositional systems at basin scale. The calcareous nanofossil assemblage is largely dominated by Coccolithus pelagicus, C. formous as well as different species of Reticulofenestra, accompanied by occasional specimens of several species of Sphenolithus and Chiasmolithus. The results obtained, based on the detailed biostratigraphic study of a composite succession 2,500 m thick, allow us to characterize the main biohorizons of the middle Eocene on the basis of global biostratigraphic standards. The studied succession was deposited during the CNE11-CNE15 or NP15-NP16 biozones, within a time span of ~3.45 Myr, between 43.96 and 40.51 Ma (Lutetian/Bartonian). This age range is compatible with the existing scheme for the South-central Pyrenean Basin. The Upper Hecho Group, between the Roncal-Fiscal megaturbidite (MT-5) and the instability facies of Gracionepel, starts in the uppermost part of Zone NP15 continues into Zone NP16. The proposed biozonation provides new data for a correlation between stratigraphic events of other regional sections and facilitates the understanding of the lateral and temporal evolution of the studied systems, as well as the improvement of the general palaeogeographic framework of the basin.
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    Tafonomía y cronología de la asociación de micromamíferos de la Cueva del Estrecho (Villares del Saz, Cuenca, España)
    (2021) Domínguez García, Ángel Carmelo; Laplana, César; Sevilla, Paloma; Guisado Di Monti, Juan Carlos; Bernárdez Gómez, María José
    La Cueva del Estrecho (Villares del Saz, Cuenca) es un yacimiento arqueo-paleontológico holoceno prácticamente inédito hasta la fecha. Se han estudiado en torno a 970 restos fósiles craneales pertenecientes a 14 taxones de micromamíferos (roedores, lagomorfos, eulipotiflos y quirópteros), procedentes del paquete sedimentario situado en la entrada de la cueva. Una datación (14C-AMS) realizada sobre hueso arroja una antigüedad de 2289-2154 años cal. BP para la asociación de micromamíferos. El estudio tafonómico realizado ha puesto de manifiesto la existencia de una elevada frecuencia de elementos digeridos en la muestra en la muestra (≥ 70%) que en algunos en algunos casos alcanzan el grado extremo. Además, se han identificado fragmentos de coprolitos adheridos a numerosos restos óseos y dentarios, así como dentición digerida de gazapos. Por todo ello puede afirmarse que el origen de la asociación fue la depredación por parte de mamíferos carnívoros, probablemente mustélidos.
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    Distinguishing between Bos and Bison petrous bones. A case study: bovines from the Des-Cubierta Cave (Pinilla del Valle, Madrid)
    (2021) Galindo-Pellicena, María A.; Arsuaga, Juan L.; Laplana, César; De Gaspar, Ignacio; Álvarez-Lao, Diego; Pérez-González, Alfredo; Baquedano, Enrique
    The taxonomic identification of large bovine remains in archaeological and palaeontological sites is important in order to infer the palaeoenvironment of these sites and to know if their inhabitants were hunters of Bosor Bison. Their presence may also have biostratigraphic or archaeozoological implications. Although the petrous bone is one of the elements of the skeleton with the greatest preservation potential in prehistoric sites, due to its hardness and compactness, it is not frequently used by the palaeontologists to distinguish between Bosand Bison, the two genera commonly present at the sites during the Pleistocene. Due to the abundance of petrous bones at the Late Pleistocene layers of the Des-Cubierta cave, the aim of this work is to identify Bosand Bisonin this site through the morphological features defined by Guadelli (1999) for this bone and using morphometric geometrics with material of Bos taurus,Bos primigenius, and Bison priscusin order to identify the differences among petrous bones of these species.
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    Diversity of large ornithopod dinosaurs in the upper Hauterivian-lower Barremian (Lower Cretaceous) of Teruel (Spain): a morphometric approach
    (2021) Verdú, Francisco J.; Cobos, Alberto; Royo-Torres, Rafael; Alcalá, Luis
    We report some large ornithopod vertebrae from two upper Hauterivian-lower Barremian (Lower Cretaceous) localities in El Castellar (Maestrazgo Basin, Teruel,Spain).These fossils have been studied systematically as well as morphometrically using a multivariate analysis in order to analyse the diversity of the sample. In fact, principal component analysis has been demonstrated as a useful tool for establishing affinities in isolated iguanodontian vertebrae, at least when size effect is not removed from the analysed dataset. As result of this study, twolarge indeterminate styracosternans are distinguished in the sample: a large one with platycoelus anterior caudal vertebrae related to the genera Magnamanusand Iguanodon, and a middle-sized one with longer-than-high dorsal vertebrae and amphicoelus anterior caudal vertebrae related to Morelladon. Such diversity of large ornithopods observed in the upper Hauterivian-lower Barremian of the Maestrazgo Basin is similar to that previously observed in Western Cameros Basin and it demonstrates the presence of at least two different forms of styracosternan in this stratigraphic range of the Iberian Peninsula.
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    Pre-Messinian ecological diversity of Mediterranean sharks revealed by the study of their dermal denticles
    (2021) Ferrón, Humberto G.; Herraiz, Jose Luis; Botella, Héctor; Martínez-Pérez, Carlos
    The Messinian salinity crisis (~5.59 Ma) is regarded as one of the most determining events for the recent evolutionary and palaeobiogeographic history of the Mediterranean biota. The impact of such episode has usually been assessed by evaluating the associated taxonomic turnover, however its consequences have rarely been interpreted from an ecological perspective. Here we assess the functional diversity of shark dermal denticles in a Serravallian locality from southeast Spain, providing a primary view into the pre-Messinian ecological diversity of shark communities from the Western Mediterranean. Our results reveal a high diversity of functional types of dermal denticles including dermal denticles that prevent the settlement of ectoparasites and epibiontes, abrasion resistant dermal denticles, drag reduction dermal denticles and dermal denticles with less specific, or more generalized, functions. This variety of dermal denticles supports the presence of several ecological groups of sharks such as schooling and strong swimming species and, possibly, demersal species, slow sharks of the open water bioluminescent taxa. Therefore, the application of this methodology to Pliocene and Pleistocene fossiliferous sites will be crucial for unravelling the impact of Messinian salinity crisis and other recent geological events from an ecological perspective, allowing us to understand the shark communitycompositionand diversityof the current Mediterranean Sea.
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    Gliridae from the late Oligocene of the province of Teruel (Spain)
    (2021) Freudenthal, Matthijs; Martín-Suárez, Elvira
    The Calatayud-Teruel Basin is one of the richest areas of Spain, when fossil mammals are concerned. Around the village of Montalbán many lower Oligocene fossil mammal localities are known and several upper Oligocene localities with rich micromammal faunas are present between the villages of Vivel del Río Martín and Martín del Río. Previous papers on these upper Oligocene localities were dedicated to the Cricetidae and to the zapodid Plesiosminthus, in this work we describe the fauna of Gliridae, composed of the genera Gliravus,Butseloglis, Peridyromys, Microdyromysand Paraglis. Among these the latter genus is very poorly represented. Microdyromysis of special importance because its presence in our material permits a new concept of the phylogeny of the genus.