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I studied simultaneously Biology and Philosophy at the University of Valencia (UV), obtaining a Ph.D. in Biology in 1983, and a Ph.D. in Philosophy in 1988. I was a Postdoctoral Fellow at the University of California, Davis (USA), in 1985-1986, and I have been invited Prof. in the University of California, Irvine (USA), in 1988 and 1994. After coming back from my post doctorate in the USA, in 1986, I created the Evolutionary Genetics Research Group at the Department of Genetics of the UV, where I am Prof. of Genetics since 1993, and have been its Director from 1995 to 1998. I was the promoter of the Cavanilles Institut of Biodiversity and Evolutionary Biology at the UV. I also promoted the creation of the Astrobiology Center (CSIC-INTA), the Public Health Research Center (CSISP), now integrated in the Valencian Region Foundation for the Promotion of Health (FISABIO) and the Institute for Integrative Systems Biology, a joint collaborative center between the UV and the Spanish Research Council, CSIC. Currently, I chair the Institutional Professorship between FISABIO and UV. I am author of more than five hundred publications about Genetics, Evolution and Philosophy of Biology. My most significant contributions are in the fields of Symbiosis and Bacterial Genomics. I am involved in the study of the Human and Animal Microbiome under an Evolutionary and Systems Biology perspective. I am founder member of the European Society for Evolutionary Biology, Spanish Society of Virology and Spanish Society of Evolutionary Biology, from which I was president between 2009-2017. I received the “Ciutat de Barcelona” award to Scientific Research (1996), “Diario Médico” award (2006), “Diploma of the President of the Valencian Government” for excellence in Biomedical Research (2010), “National Prize of Genetics” (2012), “Lilly Prize in Biomedical Research (2013)”, “Mexico Award in Science and Technology (2015)”, “Doctor Honoris Causa by Menendez Pelayo International University (UIMP, Santander, 2016). I am a fellow of the American Association for the Advancement of Science since 1998.